Paul, the 4500 BP dating, so around 2500 BCE, refers to the Mota Man skeleton from Ethiopia which has purely 'African' DNA and the 3000 BP dating, around 1000 BCE, is the proposed date for the hypothesised return migration to East Africa from the near east and so the introduction of the 'Eurasian' DNA.
The second article suggests that this migration was prompted by problems within the Egyptian Empire and the date, c. 1000 BCE ,would be during the third Intermediate Period when there were certainly significant political upheavals and weakness, droughts and all sorts of instability there, and then around the start of the first millennium CE, the ancestors of the Khoisan people began moving south.